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Four new species of Heth Cobb, 1898 (Rhigonematida: Hethidae) from Vietnamese diplopods

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Four new species of Heth are described from diplopods collected in different parts of Viet Nam: Heth vietnamensis sp. n. and H. tonkinensis sp. n. in Ba Vi National Park, Ha Tay Province; H. taybaci sp. n. in Lai Chau Province and H. taynguyeni sp. n. in Chu Yang Sin National Park, Dak Lak Province. The new species are morphologically related to those described from Papua New Guinea, but are differentiated by the form of the female cuticular armature. Heth vietnamensis sp. n. is characterised by the posterior end of the lappets overhanging the anterior pair of the lateral spines, the similar size of spines in both pairs and the presence of two pairs of small papillae on the anterior lip of the cloacal aperture. Heth tonkinensis sp. n. can be differentiated from the other species by having the longest lappets, lateral spines with fused bases and the unequal-sized spicule heads. Heth taybaci sp. n. has the shortest lappets of the described Vietnamese species, has gaps between the lateral spines that are wider than the spine base and the presence of a bursa-like cuticular fold at cloacal aperture level. Heth taynguyeni sp. n. males also have a bursa-like cuticular fold but, unlike H. taybaci sp. n., this species is characterised by the bifurcate distal tips of the spicule and the undulating inner margin of the lappets.

Affiliations: 1: Centre of Parasitology, A.N. Severtsov's Institute of Ecology and Evolution, Russian Academy of Sciences, Leninskii Prospect 33, Moscow 119071, Russian Federation


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