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A multivariate analysis of geographic variation in the Trapelus agilis complex (Sauria: Agamidae)

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The "Trapelus agilis complex" encompasses three conventional taxa: T. a. agilis (Olivier), T. a. isolepis (Boulenger), and T. a. sanguinolentus (= T. sanguinolentus) (Pallas). In previous studies the author revised this complex, synonymized T. a. isolepis with T. a. agilis, and recognized four distinct subspecies within the complex: T. a. agilis from central Iranian Plateau, southern Afghanistan, and southwestern Pakistan; T. a. sanguinolentus from northeastern Iran, northern Afghanistan, and the Central Asian Republics of the former USSR as well as Daghestan; T. a. khuzistanensis from the lowland southwestern Iran; and T. a. pakistanensis from southeastern Pakistan as well as adjoining northwestern India. In order to re-evaluate this judgment and to resolve the patterns of geographic variation within and among the mentioned four taxa, cluster analysis, principal components analysis, and canonical variate analysis were employed using 18 most informative morphological characters recorded for 522 undamaged and fully-grown specimens belonging to 19 operational taxonomic units (OTUs) throughout the range. The principal components as well as canonical variate analyses identified four primary groups of individuals corresponding to the four taxa recognized by the previous studies.


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