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A first fossil record of the terrestrial crab, Geothelphusa tenuimanus (Miyake & Minei, 1965) (Decapoda, Brachyura, Potamidae) from Okinawa Island, central Ryukyus, Japan

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Fossils of the terrestrial crab, Geothelphusa tenuimanus, are described. These fossils were collected from clay within the fissures that rift the limestone of the Middle Pleistocene Ryukyu Formation on Okinawa Island, Ryukyu Islands. The geological age of the fossil G. tenuimanus was estimated as Late Pleistocene, because they occurred in association with a fossil deer (Cervus sp.), which is already extinct on Okinawa Island. This is the first fossil record of a true freshwater crab from the Ryukyu Islands. In both fossil and extant specimens of G. tenuimanus, the longer movable fingers of the chelipeds usually have a stronger curvature, but the present fossil specimens contain higher numbers of strongly curved movable fingers than found in extant specimens. This difference may be caused by the fact that no sufficiently large extant specimens have been examined. Des fossiles du crabe terrestre Geothelphusa tenuimanus, sont décrits. Ces fossiles ont été récoltés dans l'argile, à l'intérieur de fissures qui traversent le calcaire de la formation de Ryukyu du Pléistocène moyen dans l'île d'Okinawa (îles Ryukyu). L'âge géologique du fossile G. tenuimanus a été estimé au Pléistocène supérieur, car ils ont été trouvés en association avec un cerf fossile (Cervus sp.) qui est déjà éteint sur l'île d'Okinawa. C'est la première mention de la présence d'un vrai crabe d'eau douce fossile aux îles Ryukyu. Chez les spécimens fossiles comme chez les spécimens actuels de G. tenuimanus, les doigts mobiles des chélipèdes, quand ils sont plus longs, ont habituellement une plus forte courbure, mais les spécimens fossiles ont, en plus grand nombre, des doigts mobiles plus fortement recourbés que les spécimens actuels. Cette différence peut être dûe au fait que des spécimens actuels d'une taille suffisamment grande n'ont pu être examinés.


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