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Taxonomy, biogeography and phylogenetic position of the North African Ceramius maroccanus-complex (Hymenoptera: Vespidae, Masarinae)

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image of Insect Systematics & Evolution

The Ceramius maroccanus-complex is endemic to southwestern Morocco. The status of C. maroccanus (Giordani Soika 1957) and C. montanus Gusenleitner 1990 as separate species is confirmed, C. rubripes Gusenleitner 1990 stat. n. and C. gessi sp. n. are recognized as species for the first time. The four species are redescribed/described and illustrated, and their distribution and flight period are analysed. Identification keys to males and females are provided. The C. maroccanus-complex is a monophyletic group within 'Species group 7' of Richards 1962, and is probably the sister group of the C. lusitanicus-complex.

Affiliations: 1: V. Mauss, Institut für Landwirtschaftliche Zoologie und Bienenkunde, Rheinische Friedrich- Wilhelms-Universität, Melbweg 42, D-53127 Bonn, Germany (, Email:


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