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Cratomyia macrorrhyncha, a Lower Cretaceous brachyceran fossil from the Santana Formation, Brazil, representing a new species, genus and family of the Stratiomyomorpha (Diptera)

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A new fossil Brachycera (Diptera), Cratomyia macrorrhyncha gen. n. & sp. n. is described from the Santana Formation, Lower Cretaceous, situated on the Araripe Basin (Northeastern Brazil). Its remarkable features are the large body size, very long proboscis, antenna with multi-articled flagellum, robust thorax and abdomen, and wing characters like the abbreviated costal vein, distally forked veins R4+5 and M1+2, strongly arched M1, and closed cells m3 and cup. The new genus and species does not reasonably fit in any of the existing families and a new family, Cratomyiidae is proposed. The systematic position of the family is discussed. Reasons are given for not including it in the following higher taxa of Brachycera: Xylophagomorpha, Tabanomorpha, Vermileonomorpha, Asilomorpha, and Eremoneura. The Cratomyiidae are included in Stratiomyomorpha as the sister-group of Xylomyidae + Stratiomyidae, above the origin of Pantophthalmidae.

Affiliations: 1: L. A. Mazzarolo, DCBIO/Universidade Estadual de Feira de Santana, Av. Universitaria s/n, 44.031-460 Feira de Santana BA, Brazil; 2: D. S. Amorim, Depto. de Biologia - FFCLRP/USP, Av. Bandeirantes 3900, 14.040-901 Ribei- rao Preto SP, Brazil (, Email:


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