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Geographic variation in Graphoderus zonatus (Coleoptera: Dytiscidae) in Sweden

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image of Insect Systematics & Evolution

From an analyzis of the geographic variation in Graphoderus zonatus (Hoppe), it is concluded that in Sweden there are 2 different subspecies. The southeastern, nominal subspecies, G. z. zonatus (Hoppe), is characterized by the presence of only the normal, smooth female form, and in males by a lower number (29-46) of adhesive discs on the pallette of protarsus, and by a smaller mean size. In the northern subspecies, G. z. verrucifer (Sahlberg), females are dimorphic, including also a form with granulate elytra, males have a higher number of adhesive discs (36-97), and the mean size is larger. Distributions of the 2 subspecies have been established from the principle that all provincial records including granulate females and/or males with more than 50 protarsal adhesive discs belong to G. z. verrucifer. A lectotype is designated for G. piciventris Thomson. A total of 110 males and 120 females from 21 different Swedish provinces were studied.

Affiliations: 1: A.N. Nilsson, Department of Animal Ecology, University of UmeÅ, S-90187 UmeÅ, Sweden


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