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The southern Palearctic genus Enneastigma Stein (Diptera: Anthomyiidae): Taxonomy of Mediterranean species, phylogeny and ancestral areas analysis

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Enneastigma is a small genus of myopinine anthomyiids which comprises five species in the Mediterranean subregion (E. compressum Stein, E. rarum sp. n., E. rufitibia sp. n., E. triplex Loew, E. triseriatum Stein) and two species in SE Palearctic part of China (E. pilosiventrosum Fan & Chen, E. shanghaiensis Fan & Chen). All species appear to be univoltine and only active during the cooler parts of the season. The larvae may all be coprophagous. The Mediterranean species are revised with keys to males and females and illustrated descriptions of males, females and eggs. One lectotype is designated. A cladistic analysis was conducted which also included the Chinese species. This resulted in a single, most parsimonious phylogeny. The resulting area cladogram was used to assess the alternative methods for estimating ancestral areas as proposed recently by K. Bremer and F. Ronquist. It is concluded that the best results are obtained by using a refined Ronquist analysis.

Affiliations: 1: Department of Entomology, Zoological Museum, Universitetsparken 15, 2100 Copenhagen, Denmark


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