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A revision of Neurobezzia Wirth & Ratanaworabhan and Neurohelea Kieffer, with a description of a new genus and discussion of their phylogenetic relationships (Diptera: Ceratopogonidae)

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The genus Neurohelea Kieffer is restricted to its type species, N. luteitarsis (Waltl) and is returned to the Heteromyiini. The Nearctic species previously recognized as Neurohelea nigra Wirth and N. macroneura (Malloch) are members of the Ceratopogonini and are placed in the new genus Atyphohelea. Neurohelea nigra Wirth is a new synonym of the type species Atyphohelea macroneura (Malloch), new combination. The Baltic amber species Neurohelea cothurnata (Meunier) is also placed in Atyphohelea as a new combination. An unnamed species from Taiwan is described and the pupa of A. macroneura is newly described. The male of Neurobezzia granulosa (Wirth) is described for the first time. The male previously identified as N. granulosa by Grogan & Wirth (1978) is probably the male of Amerohelea frontispina (Dow & Turner) in the Palpomyiini. Medeobezzia Yu is regarded as a new synonym of Neurobezzia Wirth & Ratanaworabhan, which now includes three species. A new world key to the genera of male and female Heteromyiini is provided. The phylogenetic relationships of the described genera are discussed.

Affiliations: 1: Research Associate, Royal British Columbia Museum and the American Museum of Natural History, 1171 Mallory Road, R1-S20-C43, Enderby, British Columbia, VOE 1V0, Canada., Email:


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