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The earliest and most oriental Calvertiellidae unearthed (Palaeodictyoptera; Late Carboniferous; China)

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We report the discovery of a new Calvertiellidae from the Xiaheyan locality (Namurian, early Late Carboniferous; Tupo Formation; China), namely Xiaheyanella ortagen. et sp. n. Besides representing both the most ancient and the most oriental occurrences for the family, the new species exhibits a derived wing venation pattern, especially if compared to the putative palaeodictyopteran cousins of the family. This discovery shows that, by the early Late Carboniferous, many insect lineages of moderate importance (i.e., considered at the familial level or below) already had a widespread distribution, and that derived conditions already accumulated. This concurs with the view that insects had already experienced a long evolutionary history by that time, remaining to be documented.

Affiliations: 1: aCollege of Life Sciences, Capital Normal University, Xisanhuangbeilu 105, Haidian District, Beijing 100048, P.R. China; 2: bCentre de Recherches sur la Paléobiodiversité et les Paléoenvironnements (CR2P, UMR 7207), Sorbonne Universités-MNHN, CNRS, UPMC-Paris 6, Muséum national d’histoire naturelle, 57 rue Cuvier, CP38, F-75005, Paris, France; 3: cGeoscience Museum, Shijiazhuang University of Economics, Huai’andonglu 136, Shijiazhuang 050031, P.R. China


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