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Two New Families of Copepods (Copepoda: Siphonostomatoida) Parasitic on Echinoderms

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Abstract The description of Dichelina phormosomae is supplemented by study of new material from a deep-water echinoid collected in the North Atlantic, and the homology of some mouthparts is reinterpreted. The phylogenetic relationships between Dichelina and other echinoderm-inhabiting siphonostomatoids are analysed. Based on inferences from the phylogenetic analysis, two new families of siphonostomatoid copepods parasitic on echinoderms are established, the Dichelinidae and the Codobidae. The hitherto unplaced genus Stephopontius is formally assigned to the family Nanaspididae.

Affiliations: 1: a The Natural History Museum, Cromwell Road, London SW7 5BD, UK ( ; 2: b Fisheries Laboratory, Hiroshima University, Minato-machi 5-8-1, Takehara, Hiroshima 725-0024, Japan (


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