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Naturally Occurring Periderm Tubes Around Secondary Phloem Fibres in the Bark of Vitis Vinifera L.

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The three-dimensional structure of periderm tubes, which normally occur in the bark of intact Vitis vinifera L. stems, is described. Each tube, which possesses several layers of mature phellem cells, differentiated around an individual strand of secondary phloem fibres within the bark. Following these peridermenclosed strands for their full length revealed that at one or two points the strand passed through a sheet of periderm and was exposed to the surrounding atmosphere. The tubes of suberised phellern within the bark isolated these strands of air-dried fibres from the living parenchyma cells around them and thus protected the bark from desiccation and pathogen attack.


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