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Revision of the Genus Titanocarcinus (Decapoda: Brachyura: Xanthoidea) with Two New Genera and One New Species

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Abstract The brachyuran genus Titanocarcinus A. Milne-Edwards, 1864, is rediagnosed and restricted to six species. It is referred to the Tumidocarcinidae Schweitzer, 2005, based upon characters of the sternum, male pleon, and dorsal carapace, along with the closely related Lobonotus A. Milne-Edwards, 1864. Several species that had been referred to Titanocarcinus are herein referred to other genera, including two new ones, Nitotacarcinus and Lathahypossia, or to other families in indeterminate genera. One new species is described from the lowermost Eocene of Spain, Titanocarcinus decor. Titanocarcinus as currently defined ranged from the Cretaceous to Eocene in northern and central Europe. Lobonotus is known only from the Eocene of North and Central America.

Affiliations: 1: a (CES, correspondence) Department of Geology, Kent State University Stark Campus, 6000 Frank Avenue NW, Canton, Ohio 44720, U.S.A. ( ; 2: b (PA) Museo Geológico del Seminario de Barcelona, Diputación 231, E-08007 Barcelona, Spain ( ; 3: c (BvB) c/o Oertijdmuseum De Groene Poort, Bosscheweg 80, NL-5283 WB Boxtel, The Netherlands ( ; 4: d (JWMJ) Natuurhistorisch Museum Maastricht, de Bosquetplein 6-7, NL-6211 KJ Maastricht, The Netherlands ( ; 5: e (HK) Mizunami Fossil Museum, Yamanouchi, Akeyo, Mizunami, Gifu 509-6132, Japan (


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